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The tests I have done : inverting the strings invert the issue, so the issue is definitly the string. I will buy another set and see.

It is flatter up the neck, but only on one string, this is the weird thing.

Yes they are wounded.
I will check the nut and the bridge.

Hello there,
I have a strange issue since I have put new strings (Optima) on my beloved MD305. The A strings are not well in tune together, a weird intonation when I go up the fret. All the others...

Just ordered one. This will be my first one, so will not be able to compare.